ClosedDensiPOPd Web Interactive: Software to investigate the impacts to density dependence by lead in bald eagles in hypothetically closed systems

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The ClosedDensiPOPd software integrates veterinary data with a population matrix model to compare the real and hypothetical impacts of lead on a population of bald eagles in a hypothetically closed system during the early and later windows of population recovery. The early window of recovery is defined as the period when abundances were low, while the late window of recovery is defined as the period when abundances were high. A closed system does not allow for incoming nor outgoing dispersal, but it does allow for temporary migration. The ClosedDensiPOPd app allows the user to compare demographic properties between lead and non-lead conditions, between time windows, between the sexes, or between all three. Comparative demographic quantities include the annual abundances, bi-annual abundances, migration and dispersal, growth rates (asymptotic, transient, cumulative, and stochastic), stable stage distribution, reproductive value, survival rates, sensitivities, elasticities, damping ratios, convergence times, and more. The ClosedDensiPOPd app is programmed to compare a population of bald eagles with and without lead toxicosis, but the software may be modified to assess the population scale impact of other veterinary maladies. For similar analyses given the full time window of population recovery (1990-2018), please use the ClosedCounterPOPd app (

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This software is being shared under a MIT license. Permission is hereby granted, free of charge, to any person obtaining a copy of this software and associated documentation files (the "Software"), to deal in the Software without restriction, including without limitation the rights to use, copy, modify, merge, publish, distribute, sublicense, and/or sell copies of the Software, and to permit persons to whom the Software is furnished to do so, subject to the following conditions: The above copyright notice and this permission notice shall be included in all copies or substantial portions of the Software. THE SOFTWARE IS PROVIDED "AS IS", WITHOUT WARRANTY OF ANY KIND, EXPRESS OR IMPLIED, INCLUDING BUT NOT LIMITED TO THE WARRANTIES OF MERCHANTABILITY, FITNESS FOR A PARTICULAR PURPOSE AND NONINFRINGEMENT. IN NO EVENT SHALL THE AUTHORS OR COPYRIGHT HOLDERS BE LIABLE FOR ANY CLAIM, DAMAGES OR OTHER LIABILITY, WHETHER IN AN ACTION OF CONTRACT, TORT OR OTHERWISE, ARISING FROM, OUT OF OR IN CONNECTION WITH THE SOFTWARE OR THE USE OR OTHER DEALINGS IN THE SOFTWARE.


This study was funded in part by the Morris Animal Foundation under Grant # D18ZO-103. This software has not been reviewed nor endorsed by the Foundation, and the views expressed in this software do not necessarily reflect the views of the Foundation, its officers, directors, affiliates or agents.

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Lefkovitch matrix; recovery; resource management


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Hanley, B. J., Dhondt, A. A., Forzán, M. J., Bunting, E. M., Pokras, M. A., Hynes, K. P., Dominguez-Villegas, E., and Schuler, K. L.. 2022. Environmental lead reduces the resilience of bald eagle populations. Journal of Wildlife Management 1– 18.

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Now showing 1 - 2 of 2
2020-12-04 20:33:13
Naming modifications and package dependencies version updates.
2019-12-09 13:58:50
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